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L. fortunei has aggressively invaded South American freshwaters, having travelled more than 5,000 km upstream across five countries. As is shown in Table 1, the main basins colonized by L. fortunei in South America are Río de la Plata basin, Guaíba basin, Patos-Mirim basin, Tramandaí basin, and Mar Chiquita basin currently. The countries colonized by L. fortunei in South America are Argentina, Brazil, Paraguay, Uruguay, and Bolivia. It is time to take some measures in advance to prevent it from invading the Amazon hydrographic basin.

Table 2

Biological traits of Limnoperna fortunei

Biological traitsMain findings
Development Main stages Nonshelled: from fertilization until the formation of the trochophore larva Planktonic larval stage is key biological trait of Limnoperna fortunei in their lifecycle. 
Shelled forms: from veliger to plantigrade larvae 
Shell Prodissoconch I: Free-swimming larval stage, reaching a length of about 115 μ
Prodissoconch II (the veliger): Free-swimming larval stage, reaching a length of about 320 μ
Nepioconch: Plantigrade stage, reaching a shell length of approximately 1,300 μm 
Dissoconch: Final shell stage, formed by the juvenile individual and becomes the permanent shell of the adult. The usual size ranges from 20 to 30 mm, some can reach about 45 mm 
Longevity Variable: 2–5 years, even over 10 years 
Diets Phytoplankton Cyanobacteria, Chlorophyceae, Xanthophyta, Bacillariophyceae, Euglenophyta, and Dinophyta Limnoperna fortunei showed a positive food selectivity for organisms with limited escape ability and low to moderate size. 
Zooplankton Rotifera, Cladocera, Copepoda, Protista, Ostracoda, and Nematoda, Limnoperna fortunei larvae 
Behaviour Great invasive potential; High reproductive rates; Strong viability It is most effective and economical to manage Limnoperna fortunei at early stages of invasion. 
Biological traitsMain findings
Development Main stages Nonshelled: from fertilization until the formation of the trochophore larva Planktonic larval stage is key biological trait of Limnoperna fortunei in their lifecycle. 
Shelled forms: from veliger to plantigrade larvae 
Shell Prodissoconch I: Free-swimming larval stage, reaching a length of about 115 μ
Prodissoconch II (the veliger): Free-swimming larval stage, reaching a length of about 320 μ
Nepioconch: Plantigrade stage, reaching a shell length of approximately 1,300 μm 
Dissoconch: Final shell stage, formed by the juvenile individual and becomes the permanent shell of the adult. The usual size ranges from 20 to 30 mm, some can reach about 45 mm 
Longevity Variable: 2–5 years, even over 10 years 
Diets Phytoplankton Cyanobacteria, Chlorophyceae, Xanthophyta, Bacillariophyceae, Euglenophyta, and Dinophyta Limnoperna fortunei showed a positive food selectivity for organisms with limited escape ability and low to moderate size. 
Zooplankton Rotifera, Cladocera, Copepoda, Protista, Ostracoda, and Nematoda, Limnoperna fortunei larvae 
Behaviour Great invasive potential; High reproductive rates; Strong viability It is most effective and economical to manage Limnoperna fortunei at early stages of invasion. 

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