The IAWPRC hypothesis, that particulate biodegradable COD hydrolyzes to a readily biodegradable (soluble) form, introduces an additional source of readily biodegradable COD for which the floc-formers and filaments compete. Acceptance of this hypothesis requires a re-evaluation of the selection criterion for the growth of filaments or floc-formers proposed by Chudoba et al. (1973). An alternative selection criterion, and some experimental observations supporting it, are presented.

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